Hey Brian;
That ‘burn’ associated with a low HR is EXACTLY what you feel when Local Muscular Endurance is the limitation to speed. It is very common to fill this in your quads when running to hiking steeply uphill outside. But the unrelenting nature of the treadmill that keeps you on your toes forces the calves into meeting their endurance limit. If you were doing this same workout on a stair machine you would not feel your calves but your quads as the limiter. If you shifted to hiking at 25% I suspect the burn will shift more to the quads.
Overall this “burn” is a good and appropriate training effect. As long as you are recovering well enough between sessions that you are seeing progress you are doing this correctly.
Keep it up.
Scott