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Hi Dan,
Thanks for writing in.
In my experience, rolling terrain has a specific effect but one need not train on it constantly. Changing of cadence, speed , and impact from rolling terrain has metabolic and signaling consequences as well as mental demands. Pacing becomes more involved to manage and the varying fatigue sensations are different than an ‘up then down’ run. If rolling terrain need be limited in use, I suggest using it for some of your intensity workouts and during the final few weeks before an ultra.
Hope this helps and best of luck.
-Seth